Reynold's genetic distance for short-term evolution (1983) [Reynolds, J. 83]
DReynold=−ln(1−θ)DReynold = -ln (1 - %theta)
the following notation is used, for all 𝚹 estimators:
nˉ=∑i=1rnir,nc=(rnˉ−∑i=1rni2rnˉ)(r−1),plu˜=∑i=1rnipilu˜rnˉ,αil˜=1−∑u=1v1pilu2˜bar n = sum _{i=1} ^r {{n_i} over r } , n_c = (r bar n - sum _{i=1} ^r {n^2 _i over {r bar n}}) over (r-1), tilde p _lu = sum _{i=1} ^r {n_i tilde p _ilu} over {r bar n},
tilde %alpha _il = 1 - sum ^{v_1} _{u=1} tilde p _ilu ^2
weighted average of single-locus ratio estimators :
θW˜=(∑l=1mal)∑l=1m(al+bl) tilde{%theta }_{W}= {( sum ^{m}_{l=1}a_{l}) } over {sum ^{m}_{l=1}( a_{l}+b_{l}) }